Subgenus Rachisoura Theobald, 1910
Tripteroides sylvestris (Theobald, 1910), original combination: Rachisoura sylvestris.
Subfamily Culicinae, genus Tripteroides. Subgenus Rachisoura includes 28 species. Subgenus abbreviation – Rac.
ADULTS ‒ Unornamented mosquitoes; vertex of head entirely or largely with dark decumbent scales, ocular scales white or whitish, 3‒6 ocular setae on either side; clypeus sometimes with scales; maxillary palpus of females 0.17‒0.30 length of proboscis, maxillary palpus of males 0.2‒1.0 length of proboscis; scutum with mainly narrow scales; acrostichal and dorsocentral setae absent; prescutellar setae reduced or absent; mesopostnotum with cluster of few minute setae (always?); postpronotal setae reduced or absent; scales of thoracic pleura white or whitish; subspiracular scales present; wing scales all broad (Filipes Group) or broad only toward apex of wing (Vanleeuweni Group); all femora with anterior surface usually entirely dark-scaled, tibiae and tarsi dark-scaled; hindungues always paired; pale scaling of abdominal terga white or whitish. FEMALE GENITALIA – Insula with 10‒35 setae in longitudinal band on either side of midline; 3 spermathecal capsules. MALE GENITALIA – Large, not retracted (fully exposed); ninth tergal lobes with slender or flattened setae; lateral plate of aedeagus with sternal lobe. LARVAE – Maxilla with short apical tooth and large articulated maxillary claw (adaptation for grasping; homologous with maxillary brush); hypostomal suture incomplete, not reaching posterior tentorial pit; seta 15-C inserted approximately midway between collar and anterior margin of labiogula; seta 6-M single, long, slender; seta 7-T strongly developed, long, multi-branched; seta 8-M,T very small, multi-branched; seta 14-I‒VI minute or absent; siphon without lateral seta. PUPAE – Seta 1-CT normally long, single, sometimes double; seta 1-I strongly developed; punctures, when present, only on terga III and IV; seta 9-VII,VIII usually well developed; paddle usually with more or less conspicuous spicules at apex. See genus Tripteroides.
The phylogenetic relationships of subgenus Rachisoura and its species have not been investigated. Belkin (1962) indicated the relationships of Rachisoura were not apparent, but noted that similarities with some species of Trichoprosopon were indicative of “genetic affinities”.
The larvae of approximately half of all species of Rachisoura are known only to inhabit Nepenthes pitchers. Other species have been found in cut bamboo, coconut shells, husks and spathes, leaf axils, tree holes and artificial containers. Larvae, as far as known, use their modified maxillae to capture and feed on other mosquito larvae that share their habitat, and perhaps feed on dead or dying insects and other arthropods as well. Females of several species are known to bite humans in the daytime, but no species of the subgenus are known to harbour or transmit pathogens of human or animal diseases.
Species of Rachisoura are found only in parts of the Australasian Region ‒ New Guinea, northern Australia and the Solomon Islands.
van den Assem, 1959 (New Guinea, taxonomy, descriptions, bionomics); Belkin, 1962 (South Pacific, Tp. mathesoni, Tp. stonei and Tp. torokinae, taxonomy, descriptions, bionomics, distributions); Mattingly, 1981 (key characters); Lee et al., 1989 (Australasian Region, keys, taxonomy, characteristics, classification, literature, distributions, bionomics); Harbach & Peyton, 1993 (comparative morphology of larval maxillae).
adentata van den Assem, 1959
bisquamatus Lee, 1946
brevirhynchus Brug, 1934
concinnus Lee, 1946
confusus Lee, 1946
cuttsi van den Assem, 1959
exnebulis Bonne-Wepster, 1948
felicitatis Bonne-Wepster, 1948
filipes (Walker, 186l)
flabelliger Bonne-Wepster, 1948
fuliginosus Lee, 1946
fuscipleura Lee, 1946
kingi Lee, 1946
latisquama (Edwards, 1927)
leei Peters, 1959
longipalpatus Lee, 1946
mathesoni Belkin, 1950
obscurus Brug, 1934
pallidus Lee, 1946
pilosus Lee, 1946
plumiger Bonne-Wepster, 1948
simplex Brug, 1934
stonei Belkin, 1950
subnudipennis (Edwards, 1927)
sylvestris (Theobald, 1910)
tityae Slooff, 1961
torokinae Belkin, 1950
vanleeuweni (Edwards, 1927)